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5/24/02 · Chargaff's Rule of Base Pairing The rules of base pairing (or nucleotide pairing) are A with T the purine adenine (A) always pairs with the pyrimidine thymine (T) C with G the pyrimidine cytosine (C) always pairs with the purine guanine (G) This is consistent with there not being enough space ( Å) for two purines to fit within the helix and too much space for two.
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